Ah yes, and I didn't show it on camera, but there was one layer I missed and spent an inordinate amount of time searching for. Turns out it was the last room in the first temple:
Two of these statues are actually teleporting statue enemies and can be killed. I missed it because the darkness of the room makes it difficult to tell them apart from the inanimate ones...
This easily-missed lair is essential to restoring the third mole hole and getting both the Ice Armor and the Shield Bracelet (the latter of which is one of the best items in the game), so be sure not to miss it!
